Tuesday, January 19, 2021

News: New Giant Titanosaur Coexisted with Giganotosaurus!

New 2021 titanosaur (MOZ Pv 1221) with other smaller sauropods from the Candeleros Formation (Otero et al., 2021. Figure 5) (Scale bar is 2 meters):

Last year, I wrote a post that talked about a possible specimen of giant sauropod (probably a titanosaur, and either closely related to, or is, Argentinosaurus) from the Candeleros Formation. Now, it seems that there is another titanosaur that coexisted with Giganotosaurus in the Candeleros Formation. It is said to have even 40 tons or more, and even been bigger than Patagotitan and probably Argentinosaurus (Otero et al., 20201, "Abstract") (Ian Randall, 2021) (Amy Woodyatt, 2021) (Bob Yirka, 2021)! The specimen is catalogued as MOZ Pv 1221.

MOZ Pv 1221 Phylogenetic Tree (Figure 4):
Discovery Site and List of Bones (Figure 1):
Scale bar length for silhouette of titanosaur is 2 meters:
Coracoid (Figure 3): 
Scale bar for coracoid is 10 cm:
Caudal (Tail) Vertebrae (Figure 2):
Scale bar for caudal vertebrae is 10 cm:
Centrum lengths for caudal vertebrae (Table 1):
I said in my post on the probable (?)Argentinosaurus specimen from Candeleros Formation that it was weird that Giganotosaurus carolinii never had giant titanosaurs to hunt within the Candeleros Formation, while it probably did hunt Patagotitan since it lived in the Albian alongside G. carolinii, but it came from a different formation. G. (Mapusaurus) roseae always hunted larger titanosaurs (Argentinosaurus and Puertasaurus, to name a few) than G. carolinii did in the Candeleros Formation (only the 80-foot long Andesaurus, but that (?)Argentinosaurus specimen is still there...). Perhaps this new unnamed giant titanosaur is related to that (?)Argentinosaurus/mystery titanosaur from Calvo (1999)? 

All I know is that G. carolinii finally has a giant titanosaur to tangle with! I will try to extrapolate a length for this animal once I can get the measurements! For now, I would say that this new titanosaur might be the same size, or perhaps bigger, than Patagotitan, just to be safe.

Size Estimation of MOZ Pv 1221:
I can't access the actual paper, but I've decided to try and formulate a size estimate for MOZ Pv 1221. 

Caudal Vertebra:
I decided to do the vertebra first. I measured the centrum length of caudal 4 (C4). I don't know how long the scale bar is, so I measured it using an estimated 5 and 10 cm each. Using 5 cm, I obtained a length of 18 cm for the centrum. Using 10 cm, I got 35 cm. I compared the centrum length of MOZ Pv 1221 to Dreadnaughtus.

1. Caudal Centrum 4:
Dreadnaughtus:
Centrum 4 Length: 33 cm.
Body Length: 30 m.

18 - 33 = 15.
15/33*100 = 45.5% decrease.
30 m - 45.5% = 54 feet (16.4 meters).

35 - 33 = 2.
2/33*100 = 6.1% increase.
30 m + 6.1% = 104 feet (31.8 meters).

Patagotitan:
Holotype's Centrum 4 Length: 36 cm (Carballido et al., 2017, Supplementary Materials, p. 32).
Body Length: 47.3 meters.

35 - 36 = 1.
1/36*100 = 2.8% decrease.
47.3 m - 2.8% = 151 feet (46.0 meters).

*Actual length: 32 cm.

Dreadnaughtus:
Centrum 4 Length: 33 cm.
Body Length: 30 m.

33 - 32 = 1.
1/33*100 = 3% decrease.
30 m - 3% = 96 feet (29.1 meters).

2. Caudal Centrum 11:
Length: 29 cm.

Dreadnaughtus:
Centrum 4 Length: 27.2 cm.
Body Length: 30 m.

27.2 - 29 = 1.8.
1.8/27.2*100 = 6.6% increase.
30 m + 6.6% = 105 feet (32.0 meters).

Caudal centrum 11 is unknown in Patagotitan.

Coracoid:
I measured MOZ Pv 1221's coracoid second.

5 cm: 41 cm.
*10 cm: 82 cm.

Patagotitan Holotype:

Coracoid: 61.5 cm (Carballido et al., 2017, Supplementary Materials, p. 33).

Body: 47.3 m.


41 - 61.5 = 20.5.

20.5/61.5*100 = 33.3% decrease.

47.3 m - 33.3% = 104 feet (31.6 meters).


*82 - 61.5 = 20.5

20.5/61.5*100 = 33.3% increase.

47.3 m + 33.3% = 207 feet (63.1 meters).

Dreadnaughtus:
Coracoid Length: 68 cm (Left).
Body Length: 30 m.

41 - 68 = 27.
27/68*100 = 39.7% decrease.
30 m - 39.7% = 59 feet (18.1 meters).

*82 - 68 = 14.
14/68*100 = 20.6% increase.
30 m + 20.6% = 119 feet (36.2 meters).

Update (2/3/21): The pubis has been stated as being 10% longer than Patagotitan's (Mercedes Benialgo, 2021, para. 4) (Madeleine Muzdakis, 2021, para. 3)

Mercedes Benialgo (2021) (Para. 4) (Translated):

Madeleine Muzdakis (2021) (Para. 3):
Only the holotype of Patagotitan (MPEF-PV 3400) has its pubis (140 cm) preserved (Carballido et al., 2017, Supplementary Materials, p. 34), and I got a length of 47.3 meters for it.

47.3 m + 10% = 171 feet (52.0 meters) for MOZ Pv 1221.
140 cm + 10% = 154 cm for MOZ Pv 1221's pubis.

If the scale bar for the silhouette of the titanosaur in the "Discovery Site and List of Bones" figure picture is 2 meters, then I've obtained a measurement of about 130 cm (at best) for the pubis. 

Update (2/9/21):
 Scale bar is 2 meters, but the pubis length is 166 cm (Discussion, para. 8):

Pubis:
Length: 166 cm.

*Patagotitan Holotype:
140 - 166 = 26.
26/140*100 = 18.6% increase.
47.3 m + 18.6% = 184 feet (56.1 meters).

*New Patagotitan pubis length: 152 cm (Otero et al., 2020, p. 9 Table 2).

152 - 166 = 14.
14/152*100 = 9.2% increase.
47.3 m + 9.2% = 170 feet (51.7 meters).

Dreadnaughtus:
Pubis: 140 cm.
Body Length: 30 cm.

140 - 166 = 26.
26/140*100 = 18.6% increase.
30 m + 18.6% = 117 feet (35.6 meters).

Futalognknosaurus:
Pubis: 137 cm (Calvo et al., 2007, p. 521).
Body Length: 31.1 meters.

137 - 166 = 29.
29/137*100 = 21.2% increase.
31.1 m + 21.2% = 124 feet (37.7 meters).

In total, MOZ Pv 1221 was 170 feet long (51.7 meters), based on the pubis compared to the holotype of Patagotitan. Based on the coracoid, a wider range of 119-207 feet (36.2-63.1 meters) was obtained. I'll go with 51.7 meters, since the authors keep using the pubis length. to give their estimate. 

As to whether or not MOZ is Argentinosaurus, the authors say that they don't know. Argentinosaurus is extremely fragmentary, so more MOZ bones will have to be excavated to see if it is a specimen of Argentinosaurus ("Discussion" para. 4):

There might be hope yet that MOZ is Argentinosaurus, and that Argentinosaurus coexisted with Giganotosaurus. However, that question will have to remain open.

Link:
Otero et al., (2021):
"Abstract":
https://www.sciencedirect.com/science/article/abs/pii/S019566712100001X
Link 2:
https://www.researchgate.net/publication/348461418_Report_of_a_giant_titanosaur_sauropod_from_the_Upper_Cretaceous_of_Neuquen_Province_Argentina
Specimen Name:
https://images.app.goo.gl/y3PyKdTeR8oA4e647
Discovery and List of Bones:
https://images.app.goo.gl/6PKmajb6s6KYZZCr5
Coracoid:
https://images.app.goo.gl/tvGwXytxtSigtrT76
Caudal (Tail) Vertebrae:
https://images.app.goo.gl/iveF5JwBh2LqRcG96
Mercedes Benialgo (2021) (Para. 4):
https://www.conicet.gov.ar/cientificos-del-conicet-hallan-un-gigantesco-dinosaurio-cuello-largo-en-neuquen/?fbclid=IwAR0jfIA8pgkgncKmN0Q9HJnV_vlYGU_tD7SpAkNLcTVgFiU885zpa0TlrAs
Madeleine Muzdakis (2021) (Para. 3): 
https://mymodernmet.com/dinosaur-fossil-titanosaur-argentina/
Ian Randall (2021):
https://www.dailymail.co.uk/sciencetech/article-9160381/Fossils-98-million-year-old-dino-unearthed-Argentina-largest-land-animal-ever.html
Amy Woodyatt (2021):
https://www.cnn.com/2021/01/19/americas/dinosaur-largest-titanosaur-intl-scli-scn/index.html
Bob Yirka (2021):
https://phys.org/news/2021-01-gigantic-dinosaur-unearthed-argentina-largest.html
My 2020 post on (?)Argentinosaurus/giant titanosaur from Candeleros Formation:
https://psdinosaurs.blogspot.com/2020/08/a-giant-sauropod-argentinosaurus-from_22.html
My lengths for the other titanosaurs mentioned in this post:
https://psdinosaurs.blogspot.com/2018/12/size-calculations-for-herbivorous.html
Lacovara et al., (2014):
https://www.researchgate.net/publication/265377777_A_Gigantic_Exceptionally_Complete_Titanosaurian_Sauropod_Dinosaur_from_Southern_Patagonia_Argentina
Link 2:
https://www.nature.com/articles/srep06196
Supplementary Materials:
https://static-content.springer.com/esm/art%3A10.1038%2Fsrep06196/MediaObjects/41598_2014_BFsrep06196_MOESM1_ESM.pdf
Carballido et al., (2017):
http://rspb.royalsocietypublishing.org/content/284/1860/20171219
Supplementary Materials (Pg. 32-34):
https://figshare.com/articles/Electronic_Supplementary_material_from_A_new_giant_titanosaur_sheds_light_on_body_mass_evolution_among_sauropod_dinosaurs/5297611
Otero et al., (2020) (P. 9 Table 2):
https://www.researchgate.net/publication/344346605_The_Appendicular_Osteology_of_Patagotitan_Mayorum_Dinosauria_Sauropoda
Calvo et al., (2007) (P. 521):
https://www.researchgate.net/publication/237289209_Anatomy_of_Futalognkosaurus_dukei_Calvo_Porfiri_Gonzalez_Riga_Kellner_2007_Dinosauria_Titanosauridae_from_the_Neuquen_Group_Late_Cretaceous_Patagonia_Argentina