Friday, November 24, 2017

Dino Bios: Carnotaurus.

Carnotaurus:
Carnotaurus' 2021 redesign based on skin impressions (Phys, 2021):
Time: 72.1-66 million years ago, Maastrichtian of the late Cretaceous period (possibly longer).
Place: South America.
Size: 32 feet (9.6 meters).
Diet: Carnivore.

Let's talk about my second favorite dinosaur: Carnotaurus!

Description:
Carnotaurus is an abelisaurid from Patagonia, South America (Bonaparte et al., 1990, "Abstract") (Ezcurra and Novas, 2016, pg. 150). It lived 72-66 million years ago (Ezcurra and Novas, 2016, p. 150) (Rafael Delcourt, 2018, Figure 1) (Leanza et al., 2004, p. 71), but it has been suggested to have lived possibly from the Campanian-Maastrichtian periods (83.6-72.1 million years ago) (Hendrickx and Bell, 2021, Materials and methods, para. 2). It was originally thought that Carnotaurus was discovered in the Gorro Frillo Formation (Bonaparte et al., 1990, "Abstract"), but now it is considered to have come from the La Colonia Formation (Ezcurra and Novas, 2016, p. 150) (Hendrickx and Bell, 2021, Materials and methods, para. 2). 

Carnotaurus was 32 feet (9.6 meters) in length. It had a relatively big head with two horns above its eyes. It had arms and hands smaller than T. rex! It's arms also supported four-fingered hands. It supported two horns on the top of its eyes that faced forward.

Carnivorous theropod dinosaurs had enamel in their teeth, so they must have had lips to cover and protect their teeth (Reisz and Larson, 2016, pg. 64-66) (Blake Eligh, 2016) (Mindy Weisberger, 2016) (Emanuela Grinberg, 2016) (Phys, 2016). Therefore, Carnotaurus would have had lips covering its teeth. Interestingly, dinosaurs couldn't move its tongues (Mindy Weisberger, 2018) (ScienceDaily, 2018).

Prey:
Dinosaurs that were on the menu for Carnotaurus were the titanosaurs Saltasaurus (28-46 feet; 8.4-13.9 meters), Argyrosaurus (78 feet; 23.8 meters), and Puertasaurus; and the ornithopod Willinakaqe (24-25 feet; 7.2-7.7 meters).

Willinakaqe (Valieri et al., 2010) (Figure 2):
Top Predator:
Carnotaurus was an abelisaur, and was the top predator of its environment. The abelisaurs were subordinate to the carcharodontosaur family for most of the Cretaceous period. This is evident with Ekrixinatosaurus and Skorpiovenator being overpowered Giganotosaurus carolinii and G. (Mapusaurus) roseae. Rugops was subordinate to Carcharodontosaurus saharicus. When the carcharodontosaurs died out, the abelisaurs rose to power. This happened at the end of the Santonian. when the Campanian period began, the abelisaurs became the top predators.

Dinosaur (2000):
Carnotaurus from Dinosaur:
Carnotaurus was the main villain for one of my favorite childhood movies, Dinosaur. Originally, it was going to be T. rex, but it was changed to Carnotaurus (Donald R. Prothero, 2016, pg. 19). This explains why Carnotsaurus was so large in the film, compared to what it was in real life. Without this movie, I would have never known, or heard, of CarnotaurusCarnotaurus hunts Aladar, and Iguanodon, in the film. Carnotaurus didn't hunt Iguanodon, since Iguanodon lived in Europe and in the early Cretaceous, but it probably hunted Willinakaqe.

Links:
Original Paper:
Bonaparte et al., (1990):
2021 Redesign:
Phys (2021):
https://phys.org/news/2021-09-scientists-reveal-fossilised-skin-bull-like.html
Size:
https://psdinosaurs.blogspot.com/2018/10/calculations-for-largest-theropods.html
Time:
Ezcurra and Novas (2016) (P. 150):
https://www.researchgate.net/publication/305011083_Theropod_dinosaurs_from_Argentina
Rafael Delcourt (2018) (Figure 1):
https://www.nature.com/articles/s41598-018-28154-x
Hendrickx and Bell (2021) (Materials and methods, para. 2):
https://www.sciencedirect.com/science/article/abs/pii/S0195667121002421
International Chronostratigraphic Chart (2020):
https://stratigraphy.org/timescale/
International Commission of Stratigraphy Website:
https://stratigraphy.org/news/130
Dinosaur Movie Info:
Donald R. Prothero (2016) (Pg. 19):
https://books.google.com/books?id=y79nCwAAQBAJ&pg=PA28&lpg=PA28&dq=carcharodontosaurus+12+to+13+meters&source=bl&ots=mQcnd67tpi&sig=Litfa9DQ_hD2JEakHd2_CWX163M&hl=en&sa=X&ved=2ahUKEwiW_sOf4P7dAhXErFkKHQGSA7oQ6AEwHnoECAoQAQ#v=onepage&q=carnotaurus&f=false
Lips:
Reisz and Larson (2016) (Pg. 64-66):
https://cansvp.files.wordpress.com/2013/08/csvp-2016-abstract-book-compressed.pdf
Blake Eligh (2016):
https://www.utoronto.ca/news/did-dinosaurs-have-lips-ask-university-toronto-paleontologist
Mindy Weisberger (2016):
https://www.livescience.com/54912-did-t-rex-have-lips.html
Emanuela Grinberg (2016):
https://www.cnn.com/2016/05/22/world/dinosaur-lips-teeth-study/index.html
Phys (2016):
https://phys.org/news/2016-06-dinosaurs-lips.html
Tongue:
Mindy Weisberger (2018):
https://www.scientificamerican.com/article/t-rex-couldnt-stick-out-its-tongue/
ScienceDaily (2018):
https://www.sciencedaily.com/releases/2018/06/180620150129.htm
Strong Neck:
Ariel H. Mendez (2012):
https://bioone.org/journals/acta-palaeontologica-polonica/volume-59/issue-3/app.2011.0129/The-Cervical-Vertebrae-of-the-Late-Cretaceous-Abelisaurid-Dinosaur-Carnotaurus/10.4202/app.2011.0129.full
Riley Black (2012):
https://www.smithsonianmag.com/science-nature/carnotaurus-had-a-hefty-neck-24787083/
Other Papers:
Skull Info.:
Cerroni et al., (2021) (Supplementary Materials):
https://figshare.com/articles/dataset/The_skull_of_i_Carnotaurus_sastrei_i_Bonaparte_1985_revisited_insights_from_craniofacial_bones_palate_and_lower_jaw/12848981?file=24407432
Prey:
Saltasaurus:
https://psdinosaurs.blogspot.com/2018/12/size-calculations-for-herbivorous.html
Zurriaguz and Powell (2015) (Pg. 284):
https://www.researchgate.net/publication/271854004_New_contributions_to_the_presacral_osteology_of_Saltasaurus_loricatus_Sauropoda_Titanosauria_from_the_Upper_Cretaceous_of_northern_Argentina
Argyrosaurus
Length:
https://psdinosaurs.blogspot.com/2018/12/size-calculations-for-herbivorous.html
Time:
Mannion and Otero (2012) (Pg. 616):
https://www.researchgate.net/publication/235635155_A_Reappraisal_of_the_Late_Cretaceous_Argentinean_Sauropod_Dinosaur_Argyrosaurus_superbus_with_a_Description_of_a_New_Titanosaur_Genus
Casal et al., (2016) (Spanish) ("Abstract," Pg. 59 "Age of the Lago Colhue Huapi Formation"/"Edad de la Formacion Lago Colhue Huapi"):
https://www.researchgate.net/publication/303598984_Ordenamiento_y_caracterizacion_faunistica_del_Cretacico_Superior_del_Grupo_Chubut_Cuenca_del_Golfo_San_Jorge_Argentina
English Version:
https://webcache.googleusercontent.com/search?q=cache:Ytyd4o1h6aEJ:https://doi.org/10.4072/rbp.2016.1.05+&cd=4&hl=en&ct=clnk&gl=us
Vallati et al., (2016) ("Abstract"):
https://app.dimensions.ai/details/publication/pub.1006664406
Lamanna et al., (2019) ("Abstract"):
https://bioone.org/journals/Annals-of-Carnegie-Museum/volume-85/issue-3/007.085.0301/A-New-Peirosaurid-Crocodyliform-from-the-Upper-Cretaceous-Lago-Colhu%c3%a9/10.2992/007.085.0301.short
Ibiricu et al., (2019) ("Abstract"):
https://www.researchgate.net/publication/337834456_New_materials_and_an_overview_of_Cretaceous_vertebrates_from_the_Chubut_Group_of_the_Golfo_San_Jorge_Basin_central_Patagonia_Argentina
Size:
https://psdinosaurs.blogspot.com/2018/12/size-calculations-for-herbivorous.html
Time:
Leanza et al., (2004) (Pg. 71):
https://www.researchgate.net/publication/257047651_Cretaceous_terrestrial_beds_from_the_Neuquen_Basin_Argentina_and_their_tetrapod_assemblages
Valieri et al., (2010) (Pg. 219-220):
https://web.archive.org/web/20110903014352/http://www.macn.secyt.gov.ar/investigacion/descargas/publicaciones/revista/12/rns_vol12-2_217-231.pdf
Link 2:
http://www.scielo.org.ar/scielo.php?script=sci_arttext&pid=S1853-04002010000200006
Garcia and Salgado (2013) ("Location and geological setting"):
https://www.app.pan.pl/archive/published/app58/app20110055.pdf
Ezcurra and Novas (2016) (Pg. 150):
https://www.researchgate.net/publication/305011083_Theropod_dinosaurs_from_Argentina